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species Pulchriphyllium bioculatum (Gray, 1832)

• Common name(s):  Javanese Leaf Insect; Gray's Leaf Insect; Javanese Leaf-insect; Gray's Leaf-insect
• Images:

  specimen

• Distribution:
  •  China, India, Java, Mauritius, Peninsular Malaysia, Seychelles, Singapore, Sri Lanka, Sumatra
• Ecology:
  • Terrestrial.
• Specimen records are available.
• Citations (103):
  • Gray, G.R. 1832. In Griffith & Pidgeon. The animal kingdom arranged in conformita with its organisation by the Baron Cuvier 15:191, pl. 63:3   >> Phyllium bioculatum    urn:lsid:Phasmida.speciesfile.org:TaxonName:6994
  • Gray, G.R. 1835. Synopsis of the species of insects belonging to the family of Phasmidae 30   >> Phyllium bioculatum
  • Burmeister. 1838. Handbuch der Entomologie 2:590   >> Phyllium bioculatum
  • Audinet-Serville. 1838[1839]. Histoire naturelle des Insectes. Orthoptères 292   >> Phyllium bioculatum
  • Gray, G.R. 1843. Zoologist 1(1):117   >> Note: Female >> Phyllium bioculatum
  • Westwood. 1859. Catalogue of the orthopterous insects in the collection of the British Museum. Part I. Phasmidae. 175   >> Phyllium bioculatum
  • Stål. 1875. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar 32:105   >> Phyllium bioculatum
  • Bolívar [Y Urrutia]. 1895. Ann. Soc. entomol. Fr. 64:374   >> Phyllium bioculatum
  • Rehn, J.A.G. 1904. Proc. Acad. Nat. Sci. Philad. 56:106   >> Phyllium bioculatum
  • Kirby, W.F. 1904. A synonymic catalogue of Orthoptera. 1. Orthoptera Euplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae) 1:421   >> Pulchriphyllium bioculatum    urn:lsid:Phasmida.speciesfile.org:TaxonName:6995
  • Redtenbacher. 1906. Die Insektenfamilie der Phasmiden 1:174   >> Phyllium bioculatum
  • Giglio-Tos. 1910. Bollettino dei Musei di Zoologia ed Anatomia comparata della Royal Università di Torino 25(625):14   >> Pulchriphyllium bioculatum
  • Ferrière. 1912. In Bolívar [Y Urrutia] & Ferrière. Orthoptera, Phasmidae of the Seychelles. Trans. Linn. Soc. Lond. 15(2):294   >> Note: in Seychelles >> Phyllium bioculatum
  • Merle. 1917. La Nature 2263:93 >> Note: rearing >> Phyllium bioculatum
  • Grimpe. 1921. Zool. Jb. Syst. 41:227   >> Note: Catalepsy, parthenogenesis; other studies >> Phyllium bioculatum
  • Showalter. 1929. The National Geographic Magazine 56(1):28, fig. >> Pulchriphyllium bioculatum
  • Rehn, J.A.G. & J.W.H. Rehn. 1934[1933]. Proc. Acad. Nat. Sci. Philad. 85:411   >> Phyllium bioculatum
  • Orian. 1957. Ann. Mag. Nat. Hist. 12 10(115):517   >> Note: Mauritius, not very common >> Pulchriphyllium bioculatum
  • Vinson. 1968. The Mauritius Institute Bulletin 6(3):113 >> Note: in list of phasmids from Mauritius >> Pulchriphyllium bioculatum
  • Klante. 1969. Abhandlungen und Berichte des Naturkundemuseums Görlitz 44(7):2 >> Phyllium bioculatum
  • Sellick. 1974. Stick and Leaf Insects 27 >> Note: rearing >> Phyllium bioculatum
  • Bedford. 1978. Annual Review of Entomology 23:126   >> Note: Oviposition >> Phyllium bioculatum
  • Robinson, D. 1980. PSG Newsletter 3:2 >> Note: rearing >> Phyllium bioculatum
  • Sellick. 1980. University of London, Unpublished thesis 197   >> Note: Egg [unpublished thesis] >> Phyllium bioculatum
  • Passerin d´Entreves. 1981. Cataloghi. IV - Collezione Ortotterologiche del Museo di Zoologia de´ll Università di Torino 55   >> Pulchriphyllium bioculatum
  • Vanschuytbroeck & Cools. 1981. Bull. Inst. r. Sci. nat. Belg. 53(23):25   >> Note: in RBINS coll >> Phyllium bioculatum
  • Hausleithner. 1983. Entomol. Z. 93(19):278 >> Note: notes, figs. >> Phyllium bioculatum
  • Brock. 1985. The Phasmid Rearer’s Handbook (Stick Insects and Leaf Insects) 28, fig >> Note: Rearing >> Phyllium bioculatum
  • Carlberg. 1986. Zool. Anz. 216(1/2):6 >> Phyllium bioculatum
  • Carlberg. 1986. Biol. Zentralbl. 105:643 >> Note: Reproduction adaptations >> Phyllium bioculatum
  • Carlberg. 1987. Z. Versuchstierkd. 29:54 >> Note: Culturing phasmids for experiments >> Phyllium bioculatum
  • Carlberg. 1987. Zool. Jb. Syst. 114:47 >> Note: Ovarian structure >> Phyllium bioculatum
  • Matyot. 1988. PSG Newsletter 37:11 >> Note: in Seychelles >> Phyllium bioculatum
  • Sellick. 1988. Zoological Journal of the Linnean Society 93:280   >> Note: In checklist of eggs studied by SEM >> Phyllium bioculatum
  • Ziegler. 1989. Entomol. Z. 99(3):30 >> Note: gynandromorph >> Phyllium bioculatum
  • Flückiger & Peltier. 1989. Gespenstschrecken figs >> Note: ex. Sri Lanka >> Phyllium bioculatum
  • Matyot. 1990. Seychelles Today 7:32 >> Note: in Seychelles >> Phyllium bioculatum
  • Preston-Mafham, K. 1990. Grasshoppers and mantids of the world 82 >> Note: note on behaviour >> Phyllium bioculatum
  • Matyot. 1990. PSG Newsletter 42:4 >> Note: in the Seychelles >> Phyllium bioculatum
  • Matyot. 1992. Phasmid Studies 1(1):18   >> Phyllium bioculatum
  • Brock. 1992. Rearing and studying stick and leaf insects. The Amateur Entomologist Volume 22 13, 46, fig   >> Note: Rearing >> Phyllium bioculatum
  • Unno. 1993. Camouflage and Mimicry of Insects 12-13, fig >> Note: Camouflage and Mimicry >> Phyllium bioculatum
  • Seow-Choen, Brock & Seow-En. 1994. Malaysian Naturalist 48(1):8   >> Note: in Singapore >> Phyllium bioculatum
  • Brock. 1995. Malayan Nat. Jour. 49(2):98   >> Phyllium bioculatum
  • Schulten. 1995. DATZ 10/95:643, fig. 4 >> Note: sketches of life history >> Phyllium bioculatum
  • Schulten. 1995. Wandelnde Blätter, Stab- und Gespenstschrecken 3:108, figs., pl. 7   >> Note: rearing >> Phyllium bioculatum
  • Ziegler. 1995. Arthropoda 3(3):32 >> Note: gynandromorph >> Phyllium bioculatum
  • Woolman & Dharmasiri. 1995. Phasmid Studies 4(2):33   >> Note: foodplants in Sri Lanka >> Phyllium bioculatum
  • Hanibeltz, Y. Nakamura, Imms & Abdullah. 1995. Phasmid Studies 4(2):60   >> Note: survival of newly-hatched nymphs >> Phyllium bioculatum
  • Brock. 1995. Insect World International 1(1):25, fig >> Phyllium (Pulchriphyllium) bioculatum
  • Deschandol. 1995. Le Monde des Phasmes 31(September):20   >> Note: Mimicry >> Phyllium bioculatum
  • Seiler, C. 1996. PSG Newsletter 69:6 >> Note: rearing observations >> Phyllium bioculatum
  • Seow-Choen. 1996. Nature Malaysiana 21(3):72, figs. >> Note: notes >> Phyllium bioculatum
  • Seow-Choen, Seow-En & Seow-An. 1996. Nature Malaysiana 21(2):43   >> Note: fig. of newly-hatched nymph >> Phyllium bioculatum
  • Tay & Seow-Choen. 1996. In Turner, I.M., Diong, S.S.L. Lim & P.K.L. Ng [Ed.]. Biodiversity and the Dynamics of Ecosystems 1:183   >> Note: foodplants in Malaysia and Singapore >> Phyllium bioculatum
  • Seow-Choen. 1997. A guide to the stick and leaf insects of Singapore 139, figs.   >> Phyllium bioculatum
  • Alderton. 1997. Your first stick insect 17, fig >> Phyllium bioculatum
  • Seow-Choen. 1998. Serangga 3(1):90   >> Note: in UKM coll. >> Phyllium bioculatum
  • Brock. 1999. Stick and Leaf Insects of Peninsular Malaysia and Singapore 160, pl. 35-38, 185   >> Note: notes, egg >> Phyllium bioculatum
  • Brock. 1999. Amateur Entomologist 26:23, 41, pl. 18b   >> Note: record for shortest lived phasmid >> Phyllium bioculatum
  • Seow-Choen. 1999. In Briffett & Chew [Ed.]. State of the natural environment in Singapore 74   >> Note: in checklist of phasmids from Singapore >> Phyllium bioculatum
  • Seow-Choen. 1999[1997]. Gardens´ Bulletin, Singapore 49(2):299   >> Note: in Singapore, status rare >> Phyllium bioculatum
  • Brock. 2000. A complete guide to breeding Stick and Leaf Insects 56, figs >> Note: rearing >> Phyllium bioculatum
  • Delfosse. 2000. Le Bulletin de Phyllie 4:3 >> Note: rearing >> Phyllium bioculatum
  • Seiler, C. & et al. 2000. Phasmiden Pflege und Zucht von Gespenstschrecken, Stabschrecken und Wandelnden Blattern im Terrarium 110, figs >> Note: rearing >> Phyllium bioculatum
  • Zompro. 2000[1999]. TenDenZen Supplement 1999:61   >> Phyllium bioculatum
  • Brock. 2000. PSG Newsletter 83:9 >> Note: On a postcard >> Phyllium bioculatum
  • Bragg. 2001. Phasmids of Borneo 191, fig.   >> Phyllium bioculatum
  • Grösser. 2001. Wandelnde Blätter. Ein Katalog aller bisher beschriebenen Phylliinae-Arten und deren Eier mit drei Neubeschreibungen. 1st Edition 53, figs >> Phyllium (Pulchriphyllium) bioculatum
  • Baxter. 2002. Rearing Stick and Leaf Insects 14, 91, figs   >> Note: rearing >> Phyllium bioculatum
  • Alderton. 2002. How to Care for your Stick Insect 16, fig >> Phyllium bioculatum
  • Brock. 2003. In Gale Research Staff. Grzimek's Animal Life Encyclopedia: Insects 226, fig. 1, 231   >> Phyllium (Pulchriphyllium) bioculatum
  • Brock. 2003. Rearing and studying stick and leaf insects. The Amateur Entomologist Volume 22 42, fig. 24   >> Note: rearing >> Phyllium (Pulchriphyllium) bioculatum
  • Whiting, M.F., Bradler & Maxwell. 2003. Nature 421(6920):265   >> Phyllium bioculatum
  • Zompro. 2004. Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea) 273   >> Phyllium (Pulchriphyllium) bioculatum
  • Otte & Brock. 2005. Phasmida Species File. Catalog of Stick and Leaf Insects of the world 275   >> Phyllium (Pulchriphyllium) bioculatum
  • Seow-Choen. 2005. Phasmids of Peninsular Malaysia and Singapore 108   >> Phyllium (Pulchriphyllium) bioculatum
  • Sorpongpaisal & Thanasinchayakul. 2006. Kamphaengsean Acad. J. 4(3):22 >> Note: in northern Thailand >> Phyllium bioculatum
  • Bragg. 2007. Phasmid Studies 15(1&2):9   >> Note: Note on authorship >> Phyllium bioculatum
  • Chen, S.C. & Y.H. He. 2008. Phasmatodea of China 367   >> Phyllium (Pulchriphyllium) pulchrifolium
  • Grösser. 2008. Wandelnde Blätter. Ein Katalog aller bisher beschriebenen Phylliinae-Arten und deren Eier mit drei Neubeschreibungen. 2nd Edition. 93, figs.   >> Phyllium (Pulchriphyllium) bioculatum
  • Gerlach & Haas. 2008. Orthopteroidea of the Seychelles islands 68   >> Phyllium bioculatum
  • Hennemann, Conle, Gottardo & Bresseel. 2009. Zootaxa 2322:80   >> Note: in Bioculatum species-group; four possible subspecies suggested for Ph. (Pu.) bioculatum Gray, 1833 are: 1) Ph. bioculatum bioculatum Gray, 1833 [Peninsular Malaysia, Singapore], 2) Ph. bioculatum agathyrsus Gray, 1843 [Sri Lanka and SE-India], 3) Ph. bioculatum pulchrifolium Audinet-Serville, 1838 [Java & Sumatra] and 4) Ph. bioculatum crurifolium Audinet-Serville, 1838 [Seychelles] >> Phyllium (Pulchriphyllium) bioculatum
  • Mandal & Yadav. 2010. Records of the Zoological Survey of India Occasional Paper 318:53   >> Note: Redescribed from the literature / material in NZSI >> Phyllium (Pulchriphyllium) bioculatum
  • Narita, Pereira, Kjellberg & Kageyama. 2010. Terrestrial Arthropod Reviews 3:74   >> Note: Gynandromorphs and intersexes >> Phyllium bioculatum
  • Harman. 2011. PSG Newsletter 125:7   >> Note: PSG culture stock origin >> Phyllium bioculatum
  • Hutacharern. 2011. Checklist of Insects in Thailand Book 2 639   >> Note: Checklist of Insects in Thailand >> Phyllium bioculatum
  • Harman. 2012. PSG Newsletter 127:12-13 >> Note: PSG culture stock origin >> Phyllium bioculatum
  • Grösser. 2013. Grundwissen Wandelnde Blätter eine Einführung in die Biologie, Haltung & Zucht 46-51, figs.   >> Note: rearing >> Phyllium bioculatum
  • Bradler. 2015. Entomologie heute 27:5, fig   >> Phyllium bioculatum
  • Seow-Choen. 2016. A taxonomic guide to the stick insects of Borneo 428, figs   >> Phyllium (Pulchriphyllium) bioculatum
  • Seow-Choen. 2017. A taxonomic guide to the stick insects of Borneo Volume II 202   >> Note: removed from Borneo fauna >> Phyllium (Pulchriphyllium) bioculatum
  • Seow-Choen. 2017. A taxonomic guide to the stick insects of Singapore 153, figs   >> Phyllium (Pulchriphyllium) bioculatum
  • Cumming & Le Tirant. 2018. Faunitaxys 6(1):2   >> Phyllium (Pulchriphyllium) bioculatum
  • Cumming, Le Tirant & Teemsma. 2018. Faunitaxys 6(2):16   >> Phyllium (Pulchriphyllium) bioculatum
  • Bradler & Buckley. 2018. In Foottit & Adler [Ed.]. Insect Biodiversity: Science and Society. Volume 2 283, fig   >> Phyllium bioculatum
  • Seow-Choen. 2018. A taxonomic guide to the stick insects of Sumatra. Volume 1 666, figs   >> Note: Correction of an error in fig. in Seow-Choen 2016 >> Phyllium (Pulchriphyllium) bioculatum
  • Seow-Choen, Seow-En, Chua & Choo. 2019. Gardens’ Bulletin Singapore 71(Suppl. 1):382   >> Note: in Bukit Timah Nature Reserve, Singapore >> Phyllium (Pulchriphyllium) bioculatum
  • Seow-Choen. 2020. A taxonomic guide to the stick insects of Sumatra. Volume 2 397   >> Note: in checklist of Sumatran species >> Phyllium (Pulchriphyllium) bioculatum
  • Bank, Cumming, Yunchang Li, Henze, Le Tirant & Bradler. 2021. Communications Biology 4(932):Supp. Table 1   >> Pulchriphyllium bioculatum bioculatum    urn:lsid:Phasmida.speciesfile.org:TaxonName:523537
  • Seow-Choen. 2021. A taxonomic guide to the stick insects of Peninsular Malaysia Volume 1 899, figs   >> Pulchriphyllium bioculatum bioculatum
  • Singh, L.R.K., Chandra & D. Gupta. 2021. In Chandra, Kosygin, Raghunathan & D. Gupta [Ed.]. Faunal Diversity of Biogeographic Zones of India: North-East 228 >> Note: in checklist list of North-East India >> Phyllium (Pulchriphyllium) bioculatum
  • Brock & Büscher. 2022. Stick and leaf-insects of the world, Phasmids pl. 3-4 & 12, 425, 563, fig   >> Note: Species account; checklist of world species >> Pulchriphyllium bioculatum bioculatum
• Type specimen information:
  • Type locality: Unknown
  • Kind of type:  holotype
  • Specimen category:  Male
  • Location of type:  OUMNH Oxford

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